The skull roof has a median ridge as well as large ridges along the medial borders of the upper temporal fenestra that mark attachments for the muscle that closed the mandible. A prominent hood at the back of the skull, the nuchal crest, marks the attachment of large neck muscles. Sizable pneumatic openings on the skull top lead into small pneumatic chambers in the skull roof. The uncrushed skull cap permitted the first endocranial reconstruction in a Triassic pterosaur and shows the brain had large cerebral lobes, from which the optic lobes bulge, and small olfactory lobes.
