The toothcomb in lemurs normally consists of six teeth (four incisors and two canines), although indriids, monkey lemurs, and some sloth lemurs only have a four-tooth toothcomb due to the loss of either a canine or an incisor. Because the lower canine is either included in the toothcomb or lost, the lower dentition can be difficult to read, especially since the first premolar (P2) is often shaped like a canine (caniniform) to fill the canine's role. In folivorous (leaf-eating) lemurs, except for indriids, the upper incisors are greatly reduced or absent. Used together with the toothcomb on the mandible (lower jaw), this complex is reminiscent of an ungulate browsing pad.
