Morphologically, standard methods such as growth temperature testing using yeast extract-malt extract agar cultivation and sporulation tests on various agars were used. As far as physiological characterization goes, the novel species differed from its close relatives in that it absorbed trehalose and ethanol, while growing on ethylamine hydrogen chloride and sodium chloride. It was also unable to grow at 37 °C and absorb glucose, and could absorb sucrose, all of which are physiological aspects that differ from its close relatives.
