His work deals with the anatomy and metamorphosis of fish, frogs, salamanders and other reptiles. He was the author of research in animal embryology and comparative anatomy on the reproduction of newts and on the development of the frog: he was the first to observe and report exactly the segmentation of the egg as well as the formation of the morula and the blastopore.. Unfortunately the fact that he did not have a real academic position prevented him from having a real scientific career and getting the reputation he deserved. Moreover, despite his skill in dissecting and preparing samples, he did not leave any collections.
