All species within the subfamily Polistinae, including R. fasciata, are understood to be eusocial. The insects fall into two categories: monogynous, with a single female reproductive, and polygynous, with several. In R. fasciata, monogyny is said to be facultative. Subordinate females' reproductive capacity is only partially inhibited biologically or behaviorally suppressed. First brood females emerge and mate before overwintering. Males emerge in late May and attempt matings with workers and foundresses. It is unknown if the foundresses monopolize reproduction, and this is an important question as it defines the function of the caste system of this species. Nests are reconstructed and eggs are laid in March.
