Regarding invertebrates, the burrowing owl seems less of a generalist. It is extremely fond of termites such as Termitidae, and Orthoptera such as Conocephalinae and Copiphorinae katydids, Jerusalem crickets (Stenopelmatidae), true crickets (Gryllidae) and grasshoppers.https://sta.uwi.edu/fst/lifesciences/sites/default/files/lifesciences/images/Athene%20cunicularia%20-%20Burrowing%20Owl.pdf Bothynus and Dichotomius anaglypticus scarab beetles (Scarabaeidae) were eaten far more often than even closely related species by many burrowing owls across central São Paulo (Brazil). Similarly, it was noted that among scorpions Bothriuridae were much preferred, among spiders Lycosidae (wolf spiders), and among millipedes (Diplopoda) certain Diplocheta. Small ground beetles (Carabidae) are eaten in quantity, while larger ones are much less popular as burrowing owl food, perhaps due to the vigorous defense the large species can put up. Earthworms are also preyed upon. Burrowing owls are also known to place the fecal matter of large herbivorous mammals around the outside of their burrows to attract dung beetles, which are used to provide a steady source of food for the owls. Burrowing owls can also predate on invertebrates attracted to artificial night lighting.
