SMAD3 and SMAD4 can also form a complex with c-Fos and c-jun at the AP-1/SMAD site to regulate TGF-β -inducible transcription .
PROPN CCONJ PROPN AUX ADV VERB DET NOUN ADP NOUN CCONJ NOUN ADP DET NOUN NOUN PART VERB PROPN ADJ NOUN .