I roto i te raumati o 1944 i Cold Spring Harbor Laboratory, McClintock tīmata rangahau pūnaha i runga i te tikanga o te tauira tae mosaic o purapura witi, me te kainga tupu unstable o tenei mosaicism. tā ia te rua hou rangatira, me te taunekeneke loci iranga i huaina e ia Dissociation (Ds) me Activator (Ac). kitea e ia e kore te Dissociation noa dissociate ranei meinga te chromosome ki te pakaru, i reira ano hoki he momo o ngā pānga i runga i ngā ira pātata, i te mea i roto i te hanga hoki etahi mutations pūmau unstable. i roto i te tīmatanga o te 1948, i hanga e ia te kitenga faahiahia e taea e rua te whakakotahitanga me te Activator tu, ranei te tūranga, i runga i te chromosome. ka kite ia i te pānga o te whakawhitinga o te Acline me te Ds e te huringa o te tauira i roto i acline i runga i ngā tauira o te purapura witi i runga i ngā momo rerekē o te mosaicism. i roto i te 1950,
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