I roto i te akoranga o te mitosis, i kite ia i te mutunga o chromatids pakaru i honoa i muri i te tāruatanga chromosome. I roto i te wāhanga o te mitaphase, i hanga e te chromosomes pakaru he piriti chromatid, i pakaru nei i te wa i neke te chromatids ki te pūtau pou. I roto i te wāhanga o te mitaphase, i hanga e McClintock he piriti chromatid, i pakaru nei i roto i te interphase o te mitosis i muri, a ka i te huringa, i te huringa, ka meinga mutation nui, i taea e ia te kitea i roto i te pūtau cytogenetics. kite ia i te pakaru me te whakakotahitanga o te chromosomes i roto i te pūtau endosperm i roto i te pūtau irradiated witi. i taea hoki e ia te whakaatu e, i roto i te tahi mau tipu tipu, i roto i te University of Missouri, i reira i kore e taea e ia te waiho i te reira, no te mea i reira i reira i reira i reira i te maha atu take,
Te Whare Wānanga o Carnegie o Washington