3" style="display: none;"> A change from a purine to a pyrimidine, or vice versa, is a transversion.</span></u><span 
	 style="display: none;">RH_Transversion</span>al substitution rates, 
 while assuming that the four nucleotide frequencies are the same and that 
 rates of substitution do not vary among sites (see <u style="text-underline: double; 
														 color: #007fbf;"><span 
	 style="color: #007fbf;"><a href="Kimura_gamma_distance.htm" style="color: #007fbf;">related 
 Gamma distance</a></span>