s males, whilst 1 biases the ratio towards females).</p></li>
<li><p>\(\overline{\overline{\eta}}\): Standardized mating fitness (zero being complete fitness ineptitude, and one being regular mating skills).</p></li>
<li><p>\(\overline{\beta}\): Fecundity (average number of eggs laid).</p></li>
<li><p>\(\overline{\xi}\): Pupation success (zero being full mortality and one full pupation success).</p></li>
</ul></div>
    <div class="section level2">
    <h2 id="migration">Migration<a class="anchor" aria-labe