gies has been a powerful and widely used tool to characterize cell identities and quantify cellular/subcellular dynamics at high temporal/spatial resolution and large scale. Complementarily, transcriptional profiles represent  cellular activities. The transcriptional differences among cells can reveal different cell states, linages and subpopulations. With the development of single-cell spatial transcriptomics, we can profile morpholigiclal and transcriptonal properties from the same cell simutaneously. We 