Pteropus
Smell

Flying foxes rely heavily on their sense of smell.They have large olfactory bulbs to process scents. They use scent to locate food, for mothers to locate their pups, and for mates to locate each other. Males have enlarged androgen -sensitive sebaceous glands on their shoulders that they use for scent-marking their territories, particularly during the mating season. The secretions of these glands vary by species — of the 65 chemical compounds identified from the glands of four species, no compound was found in all species. Males also engage in " urine washing ", meaning that they coat themselves in their own urine.
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