Cerebral cortex
Layer IV

Layer IV, the internal granular layer, contains different types of stellate and pyramidal cells, and is the main target of thalamocortical afferents from thalamus type C neurons (core-type) as well as intra-hemispheric corticocortical afferents. The layers above layer IV are also referred to as supragranular layers (layers I-III), whereas the layers below are referred to as infragranular layers (layers V and VI). African elephants, cetaceans, and hippopotamus do not have a layer IV with axons which would terminate there going instead to the inner part of layer III.
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