Pteropus
Intelligence

The megabats, including flying foxes, have the greatest encephalization quotient (brain size relative to body size) of any bat family at 1.20. This value is equivalent to that of domestic dogs. Flying foxes display behaviors that indicate a reliance on long-term information storage. Though they have wide-ranging movements and cover thousands of square kilometers annually, they are consistently able to locate the same resource patches and roosts. They will visit these resource patches consistently in a strategy known as trap-lining. They can also be conditioned to perform behaviors, such as one study where spectacled flying foxes were trained to pull a lever using juice as a reinforcement. In a follow-up to the initial study, individuals who had learned to pull the lever to receive juice still did so 3.5 years later.
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