Starfish
Body wall

The body wall consists of a thin cuticle, an epidermis consisting of a single layer of cells, a thick dermis formed of connective tissue and a thin coelomic myoepithelial layer, which provides the longitudinal and circular musculature. The dermis contains an endoskeleton of calcium carbonate components known as ossicles. These are honeycombed structures composed of calcite microcrystals arranged in a lattice. They vary in form, with some bearing external granules, tubercles and spines, but most are tabular plates that fit neatly together in a tessellated manner and form the main covering of the aboral surface. Some are specialised structures such as the madreporite (the entrance to the water vascular system), pedicellariae and paxillae. Pedicellariae are compound ossicles with forceps-like jaws. They remove debris from the body surface and wave around on flexible stalks in response to physical or chemical stimuli while continually making biting movements. They often form clusters surrounding spines. Paxillae are umbrella-like structures found on starfish that live buried in sediment. The edges of adjacent paxillae meet to form a false cuticle with a water cavity beneath in which the madreporite and delicate gill structures are protected. All the ossicles, including those projecting externally, are covered by the epidermal layer.

Several groups of starfish, including Valvatida and Forcipulatida, possess pedicellariae. In Forcipulatida, such as Asterias and Pisaster, they occur in pompom -like tufts at the base of each spine, whereas in the Goniasteridae, such as Hippasteria phrygiana, the pedicellariae are scattered over the body surface. Some are thought to assist in defence, while others aid in feeding or in the removal of organisms attempting to settle on the starfish's surface. Some species like Labidiaster annulatus, Rathbunaster californicus and Novodinia antillensis use their large pedicellariae to capture small fish and crustaceans.

There may also be papulae, thin-walled protrusions of the body cavity that reach through the body wall and extend into the surrounding water. These serve a respiratory function. The structures are supported by collagen fibres set at right angles to each other and arranged in a three-dimensional web with the ossicles and papulae in the interstices. This arrangement enables both easy flexion of the arms by the starfish and the rapid onset of stiffness and rigidity required for actions performed under stress.
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