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Urination

The urethra is the vessel through which urine exits the bladder. During urination, the urethra's smooth muscle relaxes as the bladder contracts to expel urine in a pressurized stream. Following this, the urethra re-establishes muscle tone by contracting the smooth muscle layer, and the bladder returns to a relaxed, quiescent state. Urethral smooth muscle cells are mechanically coupled to each other to coordinate mechanical force and electrical signaling in an organized, unitary fashion.