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Organogenetic studies

Early on in Warming's scientific career, the morphological-organogenetic point of was the leading principle in botanical research, and he soon became one of the most prominent workers in this branch of botany. His main works from the early period are his thesis on floral development in Euphorbia and on seed plant ovules.

Warming's doctoral thesis (in Danish) dealt with ontogeny of the cyathia of Euphorbia (Euphorbiaceae).

Part of the work was published in German the year before the thesis:

His studies of pollen and anther formation i Angiosperms and on the inflorescence of Asteraceae were published in von Hanstein's Botanische Abhandlungen:

His studies on seed plant ovules were published in French as

All these works a still cited in scientific papers by scholars of botany every now and then.

Through the 1870s, Warming became much-influenced by Darwinism. The scope of his research changed. First towards understanding ontogenesis in the light of a common descent as seen in De l’Ovule, later towards plant adaptation to environmental conditions. Again, his unparalleled ability to observe plants paired with his tropical experiencewas decisive to the route he chose.