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Ecological succession

Ecological succession is generally defined as the successive occupation of a site by different plant communities. In an accreting mudflats the outer community along the sequence represents the pioneer community which is gradually replaced by the next community representing the seral stages and finally by a climax community typical of the climatic zone. Robert Scott Troup suggested that succession began in the newly accreted land created by fresh deposits of eroded soil. The pioneer vegetation on these newly accreted sites is Sonneratia, followed by Avicennia and Nypa. As the ground is elevated as a result of soil deposition, other trees make their appearance. The most prevalent, though one of the late species to appear, is Excoecaria. As the level of land rises through accretion and the land is only occasionally flooded by tides, Heritiera fomes begins to appear.