quality:correction
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Molecular phylogeny was used by Debevic et al, 2012, to demonstrate that the bees (Anthophila) arose from deep within the Crabronidae sensu lato, which was thus rendered paraphyletic. In their study, the placement of the monogeneric Heterogynaidae was uncertain. The small family Mellinidae was not included in this analysis.

Further studies by Sann et al., 2018, elevated the subfamilies (plus one tribe and one subtribe) of Crabronidae sensu lato to family status. They also recovered the placement of Heterogyna within Nyssonini and sunk Heterogynaidae. The newly erected family, Ammoplanidae, formerly a subtribe of Pemphredoninae, was recovered as the most sister family to bees.