Southeastern mixed forests grow in below the pines. Sugar maple ("Acer saccharum"), a shade tolerant tree, grows amid the oaks and hickories in the northern part of this ecoregion. Mesic forests occur in fertile, mesic, low-elevation habitats such as deep ravines and sheltered north- or east-facing slopes. Dominant trees include American beech ("Fagus grandifolia"), tulip tree ("Liriodendron tulipifera"), northern red oak, white ash ("Fraxinus americana"), black maple ("Acer nigrum"), sugar maple ("Acer saccharum"), basswood ("Tilia americana"), and bitternut hickory ("Carya cordiformis"). Understory trees include pawpaw ("Asimina triloba") and painted buckeye ("Aesculus sylvatica"). Mesic mixed hardwood forests grow on mesic uplands, ravines, lower slopes,
Nyssa sylvatica a large deep taproot when young that makes transplanting difficult. Because of this, it is fairly uncommon in cultivation and the nursery trade. Additional characteristics include: "Nyssa sylvatica" grows in various uplands and in alluvial stream bottoms from southwestern Maine and New York, to extreme southern Ontario, central Michigan, Illinois, and central Missouri, south to southern Florida, eastern Texas, and eastern Oklahoma. It also occurs locally in central and southern Mexico. Optimum development is made on lower slopes and terraces in the Southeastern United States. "Nyssa sylvatica" is found in a variety of upland and wetland habitats in its extensive