Southeastern mixed forests grow in below the pines. Sugar maple ("Acer saccharum"), a shade tolerant tree, grows amid the oaks and hickories in the northern part of this ecoregion. Mesic forests occur in fertile, mesic, low-elevation habitats such as deep ravines and sheltered north- or east-facing slopes. Dominant trees include American beech ("Fagus grandifolia"), tulip tree ("Liriodendron tulipifera"), northern red oak, white ash ("Fraxinus americana"), black maple ("Acer nigrum"), sugar maple ("Acer saccharum"), basswood ("Tilia americana"), and bitternut hickory ("Carya cordiformis"). Understory trees include pawpaw ("Asimina triloba") and painted buckeye ("Aesculus sylvatica"). Mesic mixed hardwood forests grow on mesic uplands, ravines, lower slopes,
Fagus sylvatica depends on the location: in forest areas, "F. sylvatica" grows to over , with branches being high up on the trunk. In open locations, it will become much shorter (typically ) and more massive. The leaves are alternate, simple, and entire or with a slightly crenate margin, 5–10 cm long and 3–7 cm broad, with 6–7 veins on each side of the leaf (7–10 veins in "Fagus orientalis"). When crenate, there is one point at each vein tip, never any points between the veins. The buds are long and slender, long and thick, but thicker (to ) where the buds