Onager Stallions are either solitary or live in groups of two or three. The males have been observed holding harems of females, but in other studies, the dominant stallions defend territories that attract females. Differences in behaviour and social structure likely are the result of changes in climate, vegetation cover, predation, and hunting. The social behavior of the Asian wild ass can be very different, depending on different habitats, ranges, and even threats by predators and humans. In Mongolia and Central Asia ("E. h. hemionus" and ""E. h. kulan"), a stove onager stallion can adopt harem-type social groups with several mares
Onager ancestor of all modern equids (members of the genus "Equus") lived around 5.6 (3.9–7.8) million years ago (Mya). Direct paleogenomic sequencing of a 700,000-year-old middle Pleistocene horse metapodial bone from Canada implies a more recent 4.07 Mya for the most recent common ancestor within the range of 4.0 to 4.5 Mya. The oldest divergencies are the Asian hemiones (subgenus "E. (Asinus)", including the kulan, onager, and kiang), followed by the African zebras (subgenera "E. (Dolichohippus)", and "E. (Hippotigris)"). All other modern forms including the domesticated horse (and many fossil Pliocene and Pleistocene forms) belong to the subgenus "E. (Equus)" which