Southeastern mixed forests grow in below the pines. Sugar maple ("Acer saccharum"), a shade tolerant tree, grows amid the oaks and hickories in the northern part of this ecoregion. Mesic forests occur in fertile, mesic, low-elevation habitats such as deep ravines and sheltered north- or east-facing slopes. Dominant trees include American beech ("Fagus grandifolia"), tulip tree ("Liriodendron tulipifera"), northern red oak, white ash ("Fraxinus americana"), black maple ("Acer nigrum"), sugar maple ("Acer saccharum"), basswood ("Tilia americana"), and bitternut hickory ("Carya cordiformis"). Understory trees include pawpaw ("Asimina triloba") and painted buckeye ("Aesculus sylvatica"). Mesic mixed hardwood forests grow on mesic uplands, ravines, lower slopes,
Nyssa sylvatica long stalk and a rich source or nectar for bees. They are often dioecious so a male and female tree in proximity is required to set seed, however, many trees are also polygamo-dioecious, which means they have both male and female flowers on the same tree. The fruit is a black-blue, ovoid stone fruit, about 10 mm long with a thin, oily, bitter-to-sour tasting flesh and very popular with small bird species. There are from one to three fruits together on a long slender stalk. They are a valuable energy food for birds, especially the American robin. "Nyssa sylvatica" forms