On the Origin of Species more recent study by science historian John van Wyhe has determined that the idea that Darwin delayed publication only dates back to the 1940s, and Darwin's contemporaries thought the time he took was reasonable. Darwin always finished one book before starting another. While he was researching, he told many people about his interest in transmutation without causing outrage. He firmly intended to publish, but it was not until September 1854 that he could work on it full-time. His 1846 estimate that writing his "big book" would take five years proved optimistic. An 1855 paper on the "introduction" of species, written
Genetics and the Origin of Species variation would allow populations to evolve rapidly as environmental conditions change. This book was a landmark in the evolutionary synthesis, as it presented the union of Mendelian genetics and Darwinian theory. In "Genetics and the Origin of Species", polyploidy is considered as a type of mutation. Polyploid cells have a chromosome number that is more than twice the haploid number. The effects of polyploidy between two different species causes hybridization and even greater evolution. Natural selection in an environment produces reproductive success, which benefits the species. Speciation is a process of evolution through which new biological species are formed. Dobzhansky