Reactions to On the Origin of Species one hundred million years." Copies of the second edition were advertised as ready on 24 December, in advance of official publication on 7 January 1860. The "Saturday Review" of 24 December 1859 strongly criticised the methodology of Darwin's calculations. On 3 January 1860, Darwin wrote to Hooker about it: "Some of the remarks about the lapse of years are very good, & the Reviewer gives me some good & well deserved raps,—confound it I am sorry to confess the truth. But it does not at all concern main argument." A day later, he said to Lyell "You saw I suppose
Genetics and the Origin of Species variation would allow populations to evolve rapidly as environmental conditions change. This book was a landmark in the evolutionary synthesis, as it presented the union of Mendelian genetics and Darwinian theory. In "Genetics and the Origin of Species", polyploidy is considered as a type of mutation. Polyploid cells have a chromosome number that is more than twice the haploid number. The effects of polyploidy between two different species causes hybridization and even greater evolution. Natural selection in an environment produces reproductive success, which benefits the species. Speciation is a process of evolution through which new biological species are formed. Dobzhansky