Beech and the species "F. okamotoi", proposed by the botanist Chung-Fu Shen in 1992, comprise this subgenus. The better known "Fagus" subgenus beeches are high-branching with tall, stout trunks and smooth silver-grey bark. This group includes "Fagus sylvatica", "Fagus grandifolia", "Fagus crenata", "Fagus lucida", "Fagus longipetiolata", and "Fagus hayatae". The classification of the European beech, "Fagus sylvatica" is complex, with a variety of different names proposed for different species and subspecies within this region (for example "Fagus taurica", "Fagus orientalis", and "Fagus moesica"). Research suggests that beeches in Eurasia differentiated fairly late in evolutionary history, during the Miocene. The populations in
Nyssa sylvatica Nyssa sylvatica Nyssa sylvatica, commonly known as Tupelo, Black gum, sour gum, is a medium-sized deciduous tree native to eastern North America from the coastal Northeast USA and southern Ontario south to central Florida and eastern Texas, as well as Mexico. "Nyssa sylvatica"'s genus name "(Nyssa)" refers to a Greek water nymph; the species epithet "sylvatica" refers to its woodland habitat. The species' common name tupelo is of Native American origin, coming from the Creek words "ito" ‘tree’ and "opilwa" ‘swamp’; it was in use by the mid-18th century While these trees are often known as simply "tupelo", the fuller